Murtagh and his dragon, Thorn, attack the Varden shortly after Eragon's return, but Elven spell-casters, sent by the Elven Queen Islanzadi aid Eragon and Saphira, forcing Murtagh to flee. After the fight, Roran marries Katrina. Nasuada then orders Eragon to attend the election of the new dwarf king, insisting Saphira stay behind. Among the dwarves, Eragon is reunited with Orik, who is now the chief of his clan, Dûrgrimst Ingeitum. An unsuccessful attempt to kill Eragon is found to be the work of the clan Az Sweldn rak Anhûin. Orik successfully forces Az Sweldn rak Anhûin into exile. Having won the trust of the dwarves, Orik is elected the new dwarf king. At Orik's coronation, Saphira mends Isidar Mithrim, which Arya had shattered while rescuing Eragon from Durza.
Eragon and Saphira go to Ellesméra to continue their training. There, Oromis and Glaedr reveal that Brom, not Morzan, is Eragon's father. Glaedr also reveals the source of Galbatorix's power: Galbatorix controls hundreds of Eldunarí, or the Heart of Hearts, which preserve Dragons' consciousnesses until destroyed, even after the Dragon's physical demise. Though a Dragon cannot physically exist through its Eldunarí, a dragon could still channel its energy into the bearer of the heart, and also impart its knowledge and wisdom. Eragon then visits an Elven blacksmith, Rhunön, who helps forge Eragon a new sword, which he names Brisingr, using the substance brightsteel found under the Menoa Tree. Whenever Eragon says the name Brisingr his sword ignites in blue flames, just like when Saphira breathes fire. Before Eragon and Saphira set off to return to the Varden, Oromis says that the time has come to openly oppose Galbatorix alongside Islanzadí. Glaedr gives his Eldunarí to Eragon and Saphira before they part.
Meanwhile, after marrying Katrina, Roran is sent on various missions as part of the Varden's military. One of his targets, a convoy of the empire's supply wagons, was guarded by soldiers who feel no pain, and Roran's unit suffers extreme casualties. Only nine return to the Varden, and the commander is replaced after losing his hand. Later on, on a mission to take back one of the Surdan cities, the new commander's plan almost kills the soldiers, but Roran takes over and gives new orders. Despite saving the mission, and killing one hundred and ninety-three men on his own, Roran is charged with insubordination, and is relieved of his duties. When they arrive back at the Varden, Nasuada promotes Roran to commander and sends him on another mission. When he returns he joins the siege of the city of Feinster.
Eragon and Saphira return to the Varden in the midst of the siege of Feinster. Eragon and Arya find the leader of the city, Lady Lorana, but discover that her three magicians are attempting to create a Shade. While racing to kill the magicians, Eragon passes out as Glaedr's Eldunarí shows Oromis and Murtagh fighting near the city of Gil'ead. Galbatorix takes control of Murtagh as Oromis suffers a seizure, and Eragon watches as Oromis is slain by Murtagh. Glaedr furiously charges at Thorn, who evades and kills him. Eragon regains consciousness, and he helps Arya kill the Shade, much like Arya did in Farthen Dur. After the successful siege, Eragon tells Nasuada and Arya about Oromis and Glaedr. Nasuada then tells Eragon that the Varden plan to march to Belatona, then to Dras-Leona, and finally to Urû'baen, where they plan to kill Galbatorix.
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